FAMILIA GEOTRUPIDAE Latreille, 1802
SUBFAMILIA BOLBOCERATINAE Mulsant, 1842
Tribu Athyreini Lynch Arribálzaga, 1878 / Howden & Martinez, 1963
Athyreus acuticornis Howden, 1978
Athyreus aeneus Klug, 1843
Athyreus alvarengai Howden, 1978
Athyreus anneae Howden, 1978
Athyreus bellator Westwood, 1848
Athyreus bifurcatus Macleay, 1819 [= Athyreus herculeanus Laporte de Castelnau, 1840; Athyreus rotundus Westwood, 1848; Athyreus tuberculatus Westwood, 1848]
Athyreus bilobus Howden, 1978
Athyreus brasilicus Howden, 1978
Athyreus capricornis Howden, 1978
Athyreus cyanescens Klug, 1843
Athyreus gigas Westwood, 1848
Athyreus hastifer Felsche, 1909
Athyreus hemisphaericus Boucomont, 1902 [= Athyreus canaliculatus Boucomont, 1928]
Athyreus mouraensis Howden, 2002
Athyreus nebulosus Howden, 1999
Athyreus parvus Howden, 1978
Athyreus thilliezi Boilly, 2012 (Rio de Janeiro)
Athyreus tridens Laporte de Castelnau, 1840 [= Athyreus similis Felsche, 1909] [= Athyreus villosus Boucomont, 1902]
Athyreus tuberifer Felsche, 1909
Athyreus vavini Boucomont, 1902
Athyreus zischkai Martinez, 1953
Neoathyreus (Neoathyreus) anfractus Howden, 1985
Neoathyreus (Neoathyreus) anthracinus (Klug, 1843) [= Athyreus anthracinus Klug, 1843]
Neoathyreus (Neoathyreus) arribalzagai Martinez, 1951
Neoathyreus (Neoathyreus) bidentatus (Macleay, 1819) [= Athyreus bidentatus Macleay, 1819; = Athyreus fossulatus Westwood, 1851]
Neoathyreus (Neoathyreus) brazilensis Howden, 1985
Neoathyreus (Neoathyreus) caesariatus Howden, 1985
Neoathyreus (Neoathyreus) catharinae (Bates, 1887) [= Athyreus catharinae Bates, 1887]
Neoathyreus (Neoathyreus) centromaculatus (Felsche, 1909) [= Athyreus centromaculatus Felsche, 1909]
Neoathyreus (Neoathyreus) corinthius (Klug, 1843) [= Athyreus corinthius Klug, 1843; = Athyreus violaceus Klug, 1843]
Neoathyreus (Neoathyreus) cuspinotatus Howden, 1985
Neoathyreus (Neoathyreus) excavatus (Laporte de Castelnau, 1840)
Neoathyreus (Neoathyreus) flavithorax (Arribalzaga, 1880) [= Athyreus flavithorax Arribalzaga, 1880]
Neoathyreus (Neoathyreus) goyasensis (Boucomont, 1902) [= Athyreus goyasensis Boucomont, 1902]
Neoathyreus (Neoathyreus) illotus Howden, 1985
Neoathyreus (Neoathyreus) inermis Howden, 1985
Neoathyreus (Neoathyreus) lanuginosus (Klug, 1843)
Neoathyreus (Neoathyreus) latidorsalis Howden, 1985
Neoathyreus (Neoathyreus) moraguesi Howden, 2006
Neoathyreus (Neoathyreus) obscurus Howden, 1985
Neoathyreus (Neoathyreus) politus Howden, 1985
Neoathyreus (Neoathyreus) purpureipennis (Westwood, 1848) [= Athyreus purpureipennis Westwood, 1848]
Neoathyreus (Neoathyreus) sexdentatus (Laporte de Castelnau, 1840) [= Athyreus sexdentatus Laporte de Castelnau, 1840]
Neoathyreus (Neoathyreus) tridentatus (Macleay, 1819) [= Athyreus tridentatus Macleay, 1819] [= Athyreus bicolor Laporte de Castelnau, 1840]
Neoathyreus (Neoathyreus) viridis (Boucomont, 1902) [= Athyreus viridis Boucomont, 1902]
Parathyreus bahiae (Arrow, 1913) [= Athyreus bahiae Arrow, 1913]
Parathyreus fissicollis (Arrow, 1913) [= Athyreus fissicollis Arrow, 1913]
Parathyreus lobatus Howden, 1985
Parathyreus rectus Howden, 1985
Parathyreus trituberculatus (Klug, 1843) [= Athyreus trituberculatus Klug, 1843]
Tribu Bolboceratini Mulsant, 1842
Bolbapium bigibbosum (Luederwaldt, 1929:72) [= Bolboceras bigibbosum Luederwaldt, 1929]
Bolbapium borgmeieri Martinez, 1976:539
Bolbapium caesum Klug, 1843
Bolbapium lucidulum (Klug, 1843) [= Bolboceras lucidulum Klug, 1843]
Bolbapium minutum (Luederwaldt, 1929:71) [= Bolboceras minutum Luederwaldt, 1929]
Bolbapium paralucidulum Ide & Martinez, 1993:370
Bolbapium quadrispinosum (Luederwaldt, 1929:71) [= Bolboceras quadrispinosum Luederwaldt, 1929]
Bolbapium sculpturatum (Mannerheim, 1829:44) [= Odontaeus sculpturatum Mannerheim, 1829]
Bolbapium striatopunctatum (Laporte de Castelnau, 1840) [= Bolboceras striatopunctatum Laporte de Castelnau, 1840]
Bolbapium sulcifrons Ide & Martinez, 1993:376
Pereirabolbus castaneus (Klug, 1845:46)
Zefevazia rosascostai Martinez, 1954:200
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BIBLIOGRAFÍA
Blackwelder, R.E., 1944. (Taurocerastinae, Geotrupinae) Checklist of the coleopterous insects of México, Central America, The West Indies, and South America. U.S. National Museum Bull., 185 (2):219-220
Boilly, Olivier. (Geotrupidae, Bolboceratidae). Listado de publicaciones.
Bouchard, Patrice; Yves Bousquet; Anthony E. Davies; Miguel A. Alonso-Zarazaga; John F. Lawrence; Chris H. C. Lyal; Alfred F. Newton; Chris A. M. Reid; Michael Schmitt; S. Adam Ślipiński; Andrew B. T. Smith. 2011. Family-group names in Coleoptera (Insecta).ZooKeys 88: 1–972 (april, 2011)
Lawrence, J. F. and A. F. Newton, Jr. 1995. Families and subfamilies of Coleoptera (with selected genera, notes, references and data on family-group names). pp. 779-1006 in: J. Pakaluk and S.A. Slipinski (eds.): Biology, Phylogeny, and Classification of Coleoptera: Papers Celebrating the 80th Birthday of Roy A. Crowson. Museum i Instytut Zoologii PAN, Warszawa.
Leschen, R. A. B.; Beutel, R. G. and Lawrence, J. F. (Editors); Slipinski, A. (Associate editor). 2009. Handbook of zoology. Coleoptera, Beetles, vol. 2: Morphology and Systematics (Elateroidea, Bostrichiformia, Cucujiformia partim).
TAMU (Texas A & M University). http://insects.tamu.edu/research/collection/hallan/test/Arthropoda/Insects/Coleoptera/Family/Geotrupidae.txt
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