SUDAMERICA (SOUTH AMERICA)
AMERICA CENTRAL (CENTRAL AMERICA) ªªªªª
AMERICA DEL NORTE (NORTH AMERICA) ªªªªª
FAMILIA LUCANIDAE Latreille, 1804
SUBFAMILIA Aesalinae MacLeay 1819
Tribu Aesalini MacLeay 1819
 Aesalus neotropicalis |
Aesalus smithi |
Aesalus trogoides |
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Aesalus neotropicalis Bates, 1886 ªªªªª
Aesalus smithi Bates, 1889 ªªªªª
Aesalus trogoides Albers, 1883 ªªªªª
Lucanobium guianense Paulsen 2011: 2 (ver Paulsen 2010)
Lucanobium squamosum Howden & Lawrence, 1974 (ver Paulsen 2010)
Tribu Ceratognathini Sharp 1899: 194
Hilophyllus argentinensis (Martínez, 1981: 13) (ver Paulsen 2010)
Hilophyllus martinezi Paulsen & Mondaca, 2006 (ver Paulsen 2010)
Hilophyllus penai (Martínez, 1976: 101) (ver Paulsen 2010)
Tribu Nicagini 1861: 130
Nicagus obscurus (LeConte, 1848: 86) ªªªªª (ver Paulsen 2010)
Nicagus occultus Paulsen & Smith, 2005 ªªªªª (ver Paulsen 2010)
SUBFAMILIA Syndesinae MacLeay 1819: 103
Tribu Ceruchini Jacquelin du Val & Fairmaire 1859: 4
Ceruchus piceus (Weber, 1801: 84) ªªªªª (ver Paulsen 2010)
Ceruchus punctatus LeConte, 1869 ªªªªª (ver Paulsen 2010)
Ceruchus striatus LeConte, 1859 ªªªªª (ver Paulsen 2010)
Tribu Sinodendronini Mulsant 1842: 600
Sinodendron rugosum Mannerheim, 1843 ªªªªª (ver Paulsen 2010)
Tribu Syndesini MacLeay 1819: 103
Psilodon aequinoctiale |
Psilodon gilberti |
Psilodon luki |
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Psilodon seguy |
Psilodon xerophilicum |
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Psilodon aequinoctiale (Buquet, 1840: 375)
Psilodon gilberti Boucher, 1993
Psilodon luki (Onore, Bartolozzi, and Zilioli 2011: 43)
Psilodon schuberti Perty, 1830
Psilodon seguyi (Didier, 1929: 109)
Psilodon xerophilicum Martinez & Reyes-Castillo, 1985
SUBFAMILIA Lampriminae Macleay, 1819
Tribu Lamprimini MacLeay 1819: 97
Streptocerus speciosus (Fairmaire,1850) (ver Paulsen 2010)
SUBFAMILIA Lucaninae Latreille,1804
Tribu Brasilucanini Nikolajev 1999: 171
Brasilucanus acomus Ratcliffe, 1984 (ver Paulsen 2010)
Brasilucanus alvarengai Vulcano & Pereira, 1961 (ver Paulsen 2010)
Tribu Casignetini Reid 1999: 176
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Casignetus lindei |
Casignetus spixi |
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Casignetus humboldti (Gyllenhal in Schönherr, 1817: 197)
Casignetus lindei (Boileau, 1899b: 296)
Casignetus spixi (Perty, 1830: 54)
Tribu Chiasognathini Burmeister, 1847
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Sphaenognathus armatus |
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Sphaenognathus bellicosus |
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Sphaenognathus garleppi |
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Sphaenognathus higginsi |
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Sphaenognathus lindenii |
Sphaenognathus mandibularis |
Sphaenognathus metallescens |
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Sphaenognathus peruvianus |
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Sphaenognathus pseudocurvipes |
Sphaenognathus pubescens |
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Sphaenognathus signatus |
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Sphaenognathus taschenbergi |
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Chiasognathus beneshi Lacroix, 1978 (ver Paulsen 2010)
Chiasognathus cfr. beneshi Lacroix, 1978 ***
Chiasognathus grantii (Stephens, 1832) (ver Paulsen 2010)
Chiasognathus impubis Parry, 1870 (ver Paulsen 2010)
Chiasognathus jousellinii (Reiche,1850) (ver Paulsen 2010)
Chiasognathus latreillei (Solier,1851) (ver Paulsen 2010)
Chiasognathus mniszechii Thomson, 1862: 406 (ver Paulsen 2010)
Chiasognathus cfr. mniszechii Thomson, 1862: 406 (fem) ***
Chiasognathus sombrus Paulsen y Smith, 2010: 56. (ver Paulsen 2010)
Sphaenognathus alticollis Möllenkamp, 1912 (ver Paulsen 2010)
Sphaenognathus armatus Parry, 1872
Sphaenognathus cfr. armatus Parry, 1872 ***
Sphaenognathus bellicosus Boileau, 1904
Sphaenognathus bordoni Brochier & Chalumeau, 2000 (ver Paulsen 2010)
Sphaenognathus curvipes Benesh, 1948 (ver Paulsen 2010)
Sphaenognathus feisthamelii (Guérin-Méneville, 1838: 287) (ver Paulsen 2010)
Sphaenognathus furumii Lacroix, 1990 (ver Paulsen 2010)
Sphaenognathus garleppi Boileau, 1899
Sphaenognathus gaujoni (Oberthür, 1885: 118) (ver Paulsen 2010)
Sphaenognathus giganteus Boileau, 1911 (ver Paulsen 2010)
Sphaenognathus higginsi (Parry, 1876: 174)
Sphaenognathus kolbei Kreische, 1919 (ver Paulsen 2010)
Sphaenognathus lindenii Murray, 1857
Sphaenognathus mandibularis Boileau, 1911
Sphaenognathus metallescens DeLisle, 1967
Sphaenognathus metallifer Bomans & Lacroix, 1972
Sphaenognathus monguilloni Lacroix, 1972 (ver Paulsen 2010)
Sphaenognathus nobilis Parry, 1874 (ver Paulsen 2010)
Sphaenognathus oberon Kreische, 1922 (ver Paulsen 2010)
Sphaenognathus peruvianus (Waterhouse, 1869: 18)
Sphaenognathus prionoides Buquet, 1838 (ver Paulsen 2010)
Sphaenognathus pseudocurvipes Baba 2009: 39
Sphaenognathus pubescens Waterhouse, 1873
Sphaenognathus rotundatus Lacroix, 1987 (ver Paulsen 2010)
Sphaenognathus signatus Parry, 1874
Sphaenognathus spinifer Boileau, 1904
Sphaenognathus taschenbergi Parry, 1874
Sphaenognathus cfr. taschenbergi Parry, 1874 ***
Sphaenognathus xerophilus Bartolozzi & Onore, 2006 (ver Paulsen 2010)
Tribu Lucanini Latreille 1804: 234
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Apterocyclus honoluluensis palmatus |
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Apterocyclus honoluluensis honoluluensis Waterhouse 1871: 315 (ver Paulsen 2010)
Apterocyclus honoluluensis palmatus Van Dyke 1921: 47
Apterocyclus waterhousei Sharp in Sharp & Scott 1908: 403 (ver Paulsen 2010)
Dorcus brevis (Say, 1825: 202) ªªªªª (ver Paulsen 2010)
Dorcus parallelus (Say, 1823: 248) ªªªªª (ver Paulsen 2010)
Lucanus capreolus Linnaeus, 1763 ªªªªª (ver Paulsen 2010)
Lucanus elaphus Fabricius, 1775 ªªªªª (ver Paulsen 2010)
Lucanus mazama (LeConte, 1861: 345) ªªªªª (ver Paulsen 2010)
Lucanus placidus Say, 1825 ªªªªª (ver Paulsen 2010)
Tribu Platycerini Mulsant 1842: 593
Platycerus cribripennis Van Dyke, 1928 ªªªªª (ver Paulsen 2010)
Platycerus depressus LeConte, 1850 ªªªªª (ver Paulsen 2010)
Platycerus marginalis Casey, 1897 ªªªªª (ver Paulsen 2010)
Platycerus oregonensis Westwood, 1844 ªªªªª (ver Paulsen 2010)
Platycerus virescens (Fabricius, 1775: 817) ªªªªª (ver Paulsen 2010)
Tribu Platyceroidini Paulsen and Hawks 2008: 2
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Platyceroides agassii |
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Platyceroides thoracicus |
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Platyceroides aeneus (Van Dyke, 1928: 109) ªªªªª (ver Paulsen 2010)
Platyceroides agassii (LeConte, 1861: 345) ªªªªª
Platyceroides laticollis (Casey 1914: 373) (ver Paulsen 2010)
Platyceroides latus (Fall, 1901: 291) ªªªªª (ver Paulsen 2010)
Platyceroides opacus (Fall, 1906: 393) ªªªªª (ver Paulsen 2010)
Platyceroides thoracicus (Casey, 1895: 154) ªªªªª
Platyceroides viriditinctus (Benesh, 1942: 222) ªªªªª (ver Paulsen 2010)
Platyceropsis keeni (Casey, 1895: 153) ªªªªª (ver Paulsen 2010)
Tribu Sclerostomini Benesh 1955: 97
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Aegognathus caledoniaensis |
Aegognathus confusus |
Aegognathus iridicolor |
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 Aegognathus leuthneri damasoi |
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Altitatiayus campoi |
Altitatiayus dulceae |
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Altitatiayus koikei |
Altitatiayus rotundatus |
Altitatiayus ruficollis |
Altitatiayus trifurcatus |
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Cantharolethrus buckleyi |
Cantharolethrus elongatus |
Cantharolethrus homoderoides |
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Cantharolethrus luxerii inflexus |
Cantharolethrus peruvianus |
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Charagmophorus lineatus eikoae |
 Charagmophorus lineatus lineatus |
Charagmophorus umedai |
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Incadorcus shaunai |
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Leptinopterus affinis |
Leptinopterus asketus |
Leptinopterus assimilis |
Leptinopterus atramentarius |
Leptinopterus bicolor |
Leptinopterus burmeisteri |
Leptinopterus consimilis |
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Leptinopterus elegans |
Leptinopterus erythrocnemus |
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Leptinopterus fraternus |
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Leptinopterus gracilipes |
Leptinopterus gracilis |
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Leptinopterus inaharai |
Leptinopterus mazama |
Leptinopterus melanarius |
Leptinopterus nigrotibialis |
Leptinopterus nitidus |
Leptinopterus paranensis |
Leptinopterus pellitomarginatus |
Leptinopterus puncticollis |
Leptinopterus robustus |
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Leptinopterus vestitus |
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Leptinopterus ypirangensis |
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Macrocrates bucephalus |
Metadorcinus amuelleri |
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Metadorcinus cruentus |
Metadorcinus dentifer |
Metadorcinus ditomoides |
Metadorcinus lineatus |
Metadorcinus neotragus |
Metadorcinus plagiatus |
Metadorcinus ranki |
Metadorcinus securiformis |
Metadorcinus signatipennis |
Metadorcinus sylviae |
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Metadorcinus yamauchii |
Metadorcus ebeninus |
 Metadorcus rotundatus |
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Sclerostomus bartolozzii |
Sclerostomus costatus |
Sclerostomus damasoi |
Sclerostomus delislei |
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Sclerostomus truncatus |
Sclerostomus wendyae |
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Scortizus prodigiosus |
Scortizus signatus |
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Aegognathus aguirei Arnaud & Bomans, 2007 (ver Paulsen 2010)
Aegognathus caledoniaensis Grossi et al., 2003
Aegognathus confusus Arnaud & Bomans, 2006
Aegognathus iridicolor Lüderwaldt, 1935
Aegognathus leuthneri leuthneri Van der Poll, 1886 (ver Paulsen 2010)
Aegognathus leuthneri damasoi Arnaud & Bomans, 2006
Aegognathus similis Arnaud & Bomans, 2006 (ver Paulsen 2010)
Aegognathus soulai Arnaud & Bomans, 2004 (ver Paulsen 2010)
Aegognathus spitzi Ohaus, 1929 (ver Paulsen 2010)
Aegognathus waterhousei Leuthner, 1883 (ver Paulsen 2010)
Altitatiayus campoi Grossi & Koike 2011: 2
Altitatiayus dulceae (Bomans & Arnaud, 2002: 6)
Altitatiayus godinhorum (Bomans & Arnaud, 1996: 3) (ver Paulsen 2010)
Altitatiayus koikei Grossi in Grossi & Almeida 2011: 599
Altitatiayus rotundatus (Boileau, 1902: 43)
Altitatiayus ruficollis (Lüderwaldt, 1931: 425)
Altitatiayus trifurcatus (Grossi & Racca-Filho, 2004: 1)
Andinolucanus inesae Arnaud & Bomans, 2006 (ver Paulsen 2010)
Arnaudius bomansi Grossi & Bartolozzi 2011: 388 (ver Paulsen 2010)
Arnaudius digennaroi (Arnaud et al. 2008: 4) (ver Paulsen 2010)
Arnaudius koikei (Arnaud et al. 2007: 5) (ver Paulsen 2010)
Apterodorcus bacchus (Hope, in Westwood, 1845: 26) (ver Paulsen 2010)
Apterodorcus tristis (Deyrolle in Parry, 1870: 95) (ver Paulsen 2010)
Auxicerus aethiops Jakowlew, 1900 (ver Paulsen 2010)
Auxicerus multicolor (Boileau, 1897: 248) (ver Paulsen 2010)
Auxicerus platyceps Waterhouse, 1883 (ver Paulsen 2010)
Caenolethrus varasi (Nagel, 1932: 115) (ver Paulsen 2010)
Cantharolethrus azambrei Boileau, 1897 (ver Paulsen 2010)
Cantharolethrus buckleyi Parry, 1872
Cantharolethrus elongatus Lacroix, 1982
Cantharolethrus homoderoides Kreische, 1928 ªªªªª
Cantharolethrus luxerii luxerii (Buquet, 1843: 51) (ver Paulsen 2010)
Cantharolethrus luxerii inflexus Boileau, 1899 ªªªªª
Cantharolethrus peruvianus Heller, 1918
Cantharolethrus steinheili Parry, 1875 (ver Paulsen 2010)
Charagmophorus lineatus eikoae Nagai, 2000
Charagmophorus lineatus lineatus Waterhouse, 1895
Charagmophorus umedai Nagai, 1996
Chileistomus cucullatus (Blanchard, 1842: plate 12) (ver Paulsen 2010)
Erichius brevicollis Benesh, 1956 (ver Paulsen 2010)
Erichius caelatus (Blanchard, 1842: plate 12) (ver Paulsen 2010)
Erichius calverti (Nagel, 1932: 116) (ver Paulsen 2010)
Erichius costatus Benesh, 1956 (ver Paulsen 2010)
Erichius darwinii (Hope 1841: 302) (ver Paulsen 2010)
Erichius fairmairii fairmairii (Parry, 1864: 61) (ver Paulsen 2010)
Erichius fairmairii arriguttii Weinreich, 1958 (ver Paulsen 2010)
Erichius fasciatus (Germain, 1855: 397) (ver Paulsen 2010)
Erichius femoralis (Guérin-Méneville, 1839: 303) (ver Paulsen 2010)
Erichius franzae Weinreich, 1958 (ver Paulsen 2010)
Erichius virgatus (Nagel, 1928: 290) (ver Paulsen 2010)
Erichius vittatus vittatus (Eschscholtz, 1822: 9) (ver Paulsen 2010)
Erichius vittatus magnificus Benesh, 1956 (ver Paulsen 2010)
Erichius zabulostictus Paulsen 2010: 104 (ver Paulsen 2010)
Incadorcus ashaninka Grossi 2011: 4 (ver Paulsen 2010)
Incadorcus ayamaranus Grossi 2011: 6 (ver Paulsen 2010)
Incadorcus cuzcoensis Arnaud et al. 2007: 6 (ver Paulsen 2010)
Incadorcus damasoi Arnaud & Bomans, 2006 (ver Paulsen 2010)
Incadorcus michellae Arnaud & Bomans, 2006 (ver Paulsen 2010)
Incadorcus paulseni Grossi 2011: 9 (ver Paulsen 2010)
Incadorcus shaunai Arnaud et al., 2007
Incadorcus zugeri Arnaud & Bomans, 2006 (ver Paulsen 2010)
Leptinopterus affinis Parry, 1872
Leptinopterus asketus Grossi, 2009
Leptinopterus assimilis Grossi, 2009
Leptinopterus atramentarius Nagel, 1928
Leptinopterus bicolor Lüderwaldt, 1931
Leptinopterus burmeisteri Arrow, 1943
Leptinopterus consimilis Möllenkamp, 1900
Leptinopterus constricticollis Heller, 1924 (ver Paulsen 2010)
Leptinopterus elegans Jakowleff, 1900
Leptinopterus erythrocnemus (Burmeister, 1847: 378)
Leptinopterus femoratus (Olivier, 1789: 17) (ver Paulsen 2010)
Leptinopterus fraternus Westwood, 1874
Leptinopterus fryi (Parry, 1862: 112) (ver Paulsen 2010)
Leptinopterus gracilipes Didier, 1928
Leptinopterus gracilis Boileau, 1899
Leptinopterus ibex (Billberg, 1820: 382) (ver Paulsen 2010)
Leptinopterus inaharai Lacroix, 1982
Leptinopterus mazama Kreische, 1926
Leptinopterus melanarius (Hope in Westwood, 1845: 15)
Leptinopterus nigrotibialis Lüderwaldt, 1935
Leptinopterus nitidus Lüderwaldt, 1930
Leptinopterus paranensis Parry, 1872
Leptinopterus pellitomarginatus Lüderwaldt, 1930
Leptinopterus puncticollis Lüderwaldt, 1930
Leptinopterus robustus Lüderwaldt, 1930
Leptinopterus suturalis Lüderwaldt, 1930 (ver Paulsen 2010)
Leptinopterus tibialis (Eschscholtz, 1822: 5) (ver Paulsen 2010)
Leptinopterus cfr. tibialis (Eschscholtz, 1822: 5) ***
Leptinopterus vestitus Benesh, 1937
Leptinopterus v-nigrum (Hope in Westwood, 1845: 15) (ver Paulsen 2010)
Leptinopterus ypirangensis Lüderwaldt, 1930
Leptinopterus sp. 1 ***
Leptinopterus sp. 2 ***
Leptinopterus sp. 3 ***
Leptinopterus sp. 4 ***
Macrocrates australis Lüderwaldt, 1934 (ver Paulsen 2010)
Macrocrates bucephalus (Westwood, 1845: 15)
Metadorcinus amuelleri (Weinreich, 1963: 211)
Metadorcinus auritus Kreische, 1922 (ver Paulsen 2010)
Metadorcinus beneshi (Martínez, 1953: 43)
Metadorcinus buckleyi (Waterhouse, 1886: 497) (ver Paulsen 2010)
Metadorcinus cruentus (Burmeister, 1847: 425)
Metadorcinus dentifer (Möllenkamp, 1912: 301)
Metadorcinus ditomoides (Westwood, 1855: 208)
Metadorcinus lineatus (Deyrolle, 1864: 319)
Metadorcinus neotragus (Westwood, 1855: 207)
Metadorcinus plagiatus (Burmeister, 1847: 425)
Metadorcinus ranki Grossi & Vaz-de-Mello, 2007
Metadorcinus securiformis (Lüderwaldt, 1934: 389)
Metadorcinus signatipennis (Deyrolle, 1864: 319)
Metadorcinus sylviae Boucher (1993: 420)
Metadorcinus tucumanus (Nagel, 1932: 119)
Metadorcinus yamauchii Arnaud et al., 2008
Metadorcus ebeninus Deyrolle, 1864
Metadorcus rotundatus Parry, 1862
Metadorcus sp. 1 ***
Onorelucanus aequatorianus Bartolozzi & Bomans, 1989 (ver Paulsen 2010)
Onorelucanus boileaui (Weinreich, 1960: 81) (ver Paulsen 2010)
Onorelucanus incredibilis (Arnaud et al. 2008: 3) (ver Paulsen 2010)
Onorelucanus marujae (Arnaud et al., 2008: 5) (ver Paulsen 2010)
Onorelucanus nigrosuturalis (Weinreich, 1963) (ver Paulsen 2010)
Onorelucanus noguchii (Arnaud & Bomans, 2007: 3) (ver Paulsen 2010)
Onorelucanus onorei Lacroix & Bartolozzi, 1989
Onorelucanus pulverosus (Westwood, 1875: 243) (ver Paulsen 2010)
Onorelucanus rufolineatus (Nagel, 1928) (ver Paulsen 2010)
Pycnosiphorus lessonii lessonii (Buquet, 1842: 283) (ver Paulsen 2010)
Pycnosiphorus lessonii elongatus (Deyrolle in Parry 1870: 96) (ver Paulsen 2010)
Pycnosiphorus lessonii mandibularis (Solier, 1851: 56) (ver Paulsen 2010)
Pycnosiphorus lessonii marginipennis Deyrolle in Parry, 1870 (ver Paulsen 2010)
Pycnosiphorus lessonii ugartei Paulsen 2010:122 (ver Paulsen 2010)
Pycnosiphorus philippi (Westwood, 1864: 61) (ver Paulsen 2010)
Sclerostomulus nitidus (Benesh, 1955: 99) (ver Paulsen 2010)
Sclerostomulus tuberculatus (Solier, 1851: 54) (ver Paulsen 2010)
Sclerostomus bartolozzii Arnaud & Bomans, 2007
Sclerostomus costatus (Westwood, 1845: 27)
Sclerostomus damasoi Arnaud & Bomans, 2006
Sclerostomus delislei Weinreich, 1961
Sclerostomus gounellei (Boileau, 1899b: 299) (ver Paulsen 2010)
Sclerostomus sulcicollis (Möllenkamp, 1912: 301) (ver Paulsen 2010)
Sclerostomus truncatus Lüderwaldt, 1935
Sclerostomus wendyae Arnaud & Bomans, 2006
Scortizus maculatus (Klug, 1825: 432) (ver Paulsen 2010)
Scortizus prodigiosus Arnaud et al., 2007
Scortizus signatus Weinreich, 1963
Scortizus zischkai Martínez, 1957 (ver Paulsen 2010)
Zikanius aurocinctus (Boileau, 1899b: 298) (ver Paulsen 2010)
Página creada: 13 diciembre 2009; 17:33 |
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BIBLIOGRAFÍA
Benesh, B. 1960. Coleopterorum Catalogus Supplementa, pars 8: Lucanidae. W. Junk, Gravenhage, Netherlands. 178 pp.
Blackwelder, R.E., 1944; (Lucanidae) Checklist of the coleopterous insects of México, Central America, The West Indies, and South America. U.S. National Museum Bull., 185 (2):195-197
Bouchard, Patrice; Yves Bousquet; Anthony E. Davies; Miguel A. Alonso-Zarazaga; John F. Lawrence; Chris H. C. Lyal; Alfred F. Newton; Chris A. M. Reid; Michael Schmitt; S. Adam Ślipiński; Andrew B. T. Smith. 2011. Family-group names in Coleoptera (Insecta).ZooKeys 88: 1–972 (april, 2011)
Holloway, B.A. 1960. Taxonomy and phylogeny in the Lucanidae (Insecta: Coleoptera). Records of the Dominion Museum 3: 321-365.
Holloway, B.A. 1968. The relationship of Syndesus MacLeay and Sinodendron Schneider (Coleoptera: Lucanidae). New Zealand Journal of Science 11: 264-269.
Holloway, B.A. 1969. Further studies on generic relationships in Lucanidae (Insecta: Coleoptera) with special reference to the ocular canthus. New Zealand Journal of Science 12: 958-977.
Holloway, B.A. 1998. A re-evaluation of the genera of New Zealand aesaline stag beetles (Coleoptera: Lucanidae). Journal of the Royal Society of New Zealand 28: 641-656.
Howden, H. and Lawrence, J.H. 1974. The New World Aesalinae, with notes on the North American lucanid subfamilies (Coleoptera, Lucanidae). Canadian Journal of Zoology 52: 1505-1510.
Lawrence, J. F. and A. F. Newton, Jr. 1995. Families and subfamilies of Coleoptera (with selected genera, notes, references and data on family-group names). pp. 779-1006 in: J. Pakaluk and S.A. Slipinski (eds.): Biology, Phylogeny, and Classification of Coleoptera: Papers Celebrating the 80th Birthday of Roy A. Crowson. Museum i Instytut Zoologii PAN, Warszawa.
Leschen, R. A. B.; Beutel, R. G. and Lawrence, J. F. (Editors); Slipinski, A. (Associate editor). 2009. Handbook of zoology. Coleoptera, Beetles, vol. 2: Morphology and Systematics (Elateroidea, Bostrichiformia, Cucujiformia partim).
Mizumuma, T. and S. Nagai. 1994. The Lucanid Beetles of the World. Mushi-sha, Tokyo. 337 pp.
Paulsen, M.J. 2008. Generic guide to New World Lucanidae. On-line version 2.0
Paulsen, M.J. 2010. Generic guide to New World Lucanidae. On-line version 3.0
Paulsen, M. J. 2010. The stag beetles of southern South America. Bull. U. Nebraska State Museum vol.24
Ríos, J. C. y Salazar E., J. A. 2005. Sobre algunas localidades colombianas para conocer y estudiar a Cantharolethrus Luxerii ( Buquet); Sphaenognathus hemiphanestus (Deliste)
& Sph. feisthameli (Guérin-Ménéville). (Coleóptera: Lucanidae). Boletín Científico, Centro de Museos,
Museo de Historia Natural Vol. 9, enero - diciembre, 2005, pags. 167-177.
Smith, A.B.T., 2006. A review of the family-group names for the superfamily Scarabaeoidea (Coleoptera) with corrections to nomenclature and current classification. The Coleopterist Bulletin 60: 144-204.
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